His major contribution in the field of biophysics was the demonstration of the electrical nature of the conduction of various stimuli (., wounds, chemical agents) in plants, which were earlier thought to be of a chemical nature. These claims were later proven experimentally.  He was also the first to study the action of microwaves in plant tissues and corresponding changes in the cell membrane potential. He researched the mechanism of the seasonal effect on plants, the effect of chemical inhibitors on plant stimuli and the effect of temperature. From the analysis of the variation of the cell membrane potential of plants under different circumstances, he hypothesised that plants can "feel pain, understand affection etc."
Robert malthus essay on populations
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